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mfd_shallow_16S

The scripts in this repository are part of the Microflora Danica project. The holds the workflow used to create observational tables of 16S and 18S rRNA gene fragments from raw short-read metagenomes.

Be advised, that for the metagenomic-derived data, the term "OTU" is only used due to format requirements by ampvis2, and they do not represent classical OTUs. The generated profiles can be thought of as taxonomic bins.

Scripts

Bash processing

scripts/bcl2fastq.sh performs demultiplexing using bcl2fastq based on a sample sheet per run. The sample sheet file is formatted to output data in the following format:

Folder Content
run folder A folder of the entire sequencing run.
├── lane folder/ One folder per flow cell lane
│ ├── library folder/ One folder per library ID
│ │ ├── seq files Two files (forward and reverse) per library ID

scripts/trim_rename.sh performs trimming and quality filtering of the raw data using fastp. Compresses the outputs using pigz as the internal zipping function of fastp is corrupt. The output is structured in the following format:

Folder Content
run folder A folder of the entire sequencing run.
├── lib folder/ One folder per library ID
│ ├── seq files Two files (forward and reverse) per library ID

scripts/classify_shallow_metagenomes.sh identifies the 16S and 18S using Hidden Markow Models and nhmmer, extracts the reads using awk, grep, sort, uniq and zcat, and performs taxonomic classification using SINTAX. The output is structured in the following format (simpliied to only using results for 16S):

Folder Content
run folder A folder of the entire sequencing run.
├── lib folder/ One folder per library
│ ├── forward/
│ │ ├── arc_LIB-MJXXX-A1_01_forward.hmmout.txt Output from nhmmer using arc.hmm
│ │ ├── bac_LIB-MJXXX-A1_01_forward.hmmout.txt Output from nhmmer using bac.hmm
│ │ ├── arc_bac_LIB-MJXXX-A1_01_forward.fq Extracted forward reads in fastq format
│ │ ├── arc_bac_LIB-MJXXX-A1_01_forward_MFG_ssu_database_NR987_trunc.sintax Forward SINTAX file
│ ├── reverse/
│ │ ├── arc_LIB-MJXXX-A1_01_reverse.hmmout.txt Output from nhmmer using arc.hmm
│ │ ├── bac_LIB-MJXXX-A1_01_reverse.hmmout.txt Output from nhmmer using bac.hmm
│ │ ├── arc_bac_LIB-MJXXX-A1_01_reverse.fq Extracted reverse reads in fastq format
│ │ ├── arc_bac_LIB-MJXXX-A1_01_reverse_MFG_ssu_database_NR987_trunc.sintax Reverse SINTAX file
│ ├── tmp/
│ │ ├── forward_IDs.txt Forward read IDs from nhmmer outputs
│ │ ├── forward_IDs_unique.txt Unique forward read IDs
│ │ ├── reverse_IDs.txt Reverse read IDs from nhmmer outputs
│ │ ├── reverse_IDs_unique.txt Unique reverse read IDs

scripts/find_hmm.sh finds all nhmmer output files and creates symlinks.

scripts/find_sintax.sh finds all SINTAX output files and creates symlinks.

R processing

scripts/filter_reads.R filters the combined pool of 16S rRNA gene reads to within the region between the 8F and 1391R primers for bacteria and 20F and SSU1000ArR for archaea. Splits output based on whether the origin was an MFD sample or an included control.

scripts/arcbac_sintax_to_combined_tax.R creates one big file from all SINTAX classifications. Filters the file so that either the forward or the reverse classification file is included if both are present. The read with the deepest classification is used. If the same depth is achieved the forward read is used.

scripts/arcbac_combined_tax_to_OTU.R transforms the outputs of arcbac_sintax_to_combined_tax.R to observational tables. The term "OTU" is only used for compatability with ampvis2.

To yield the final table N/A is changed to 0 using sed -i 's/N\/A/0/g' mfd_shallow_16S/R/release/DATE__MFD_arcbac_shallow_release.csv

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